Robuta

https://pmc.ncbi.nlm.nih.gov/articles/PMC34369/ The chimeric eukaryote: Origin of the nucleus from the karyomastigont in amitochondriate protists -... We present a testable model for the origin of the nucleus, the membrane-bounded organelle that defines eukaryotes. A chimeric cell evolved via symbiogenesis by... chimericeukaryoteorigin